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Mat is a young male rat that lives among a rat commune in an abandoned wine cellar, together with his best friend Toby and other rodents. He has fallen in love with Nat, a female house mouse kept as a pet on the upper floors. To get closer to her, Mat decides to become a Scavenger, allowing him to sneak up to see her more officially. Since they come from different worlds, they devise a plan to escape to the "outside" together. But Mat's overconfidence puts them in danger, and not everything in the commune is as it appears.

In a series of unlucky events, Mat finds himself alone, far away from the house. He must make his way back home, overcoming a journey across woodlands, caves, and swamps, defying overwhelming danger of a winged, poisonous, and scaled variety. With a rivet for a sword and nails as throwing daggers, Mat will brave a world to which he is unwelcome.

Help the Scavenger get back to his friends and save the rat commune from a threat that is bigger than every cheese in the world!