A stationary aeroplane on a treadmill will obviously move with the treadmill. I assume an aeroplane moving at like 1 km/h still gets pulled backward by the treadmill.

so, every wheel or ball or any other kind of rolling-thing has rolling resistance, which is how we sum up the total drag on the system. A steel ball bearing on a steel plate will have a significantly lower rolling resistance than, say, a steel cube on that same plate. Tires have some- but not a lot- of rolling resistance.

You can see that in a car, just put it into neutral and watch as you slow down, even on flat ground. Plane wheels also have rolling resistance. it's just the way our world works. But it's generally ignored because it's hard to model perfectly and in any case pretty negligible relative to the amount of acceleration being put out by modern aircraft engines.

A treadmill will only push an aircraft or whatever else along, with an acceleration that is equal to, or lower, than the rolling resistance. If you try to accelerate the plane faster, it'll 'slip', and the plane will remain largely stationary- like the dishes in the tablecloth trick (if you want to try that at home... make sure the tablecloth doesn't have a hem, heh.)

But, keep in mind you're thinking about the plane relative to either the ground, or the treadmill's belt.

the plane's wings and it's engines are 'thinking' about the plane relative to the air it's moving through. It's the airspeed that generates the lift, and the engine isn't coupled to the wheels, they're just rolling along doing their thing. (aircraft engines work by taking a volume of air and accelerating it. newton's equal-and-opposite does the rest.)